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Clank (real designation XJ0461, serial number B5429671) is the secondary titular protagonist and playable character of the Ratchet & Clank series. He is a robot created by the Zoni, who possesses a Zoni soul, and is Ratchet's best friend.

Clank was created by Orvus, the leader of the Zoni, in the Great Clock. He was produced in a blarg robot factory on Quartu which produced sentry-bots, and upon acquiring an infobot and learning about Chairman Drek's plans, he fled the planet in a stolen starship, but was shot down and crash-landed on Veldin. There, he met Ratchet, who would become his lifelong best friend afterward, as they would partake on many adventures. When the two visited the Polaris Galaxy in their mission to defeat Emperor Percival Tachyon, Clank was met by a race invisible to the others known as the Zoni, who referred to him as the Chosen One. The Zoni helped Clank defeat Tachyon and afterwards abducted him from Ratchet to take him back to the Great Clock, where he was supposed to take up his true destiny as the Senior Caretaker of the Clock with Sigmund, though after stopping Dr. Nefarious from trying to use the Clock to his advantage, Clank made Sigmund its caretaker and returned to his place at Ratchet's side.

Clank is a highly intelligent robot who is highly knowledgeable in a variety of topics and able to communicate with many other machines. He often acts as a contrast to Ratchet's more brash personality, serving as his voice of reason. Clank also helps Ratchet in the form of attachments such as the Heli-Pack allowing Ratchet to reach greater heights and storing all of Ratchet's weapons, as well as being able to access areas Ratchet cannot. Clank is also capable of commanding help, in the form of Gadge-Bots, Microbots and Zoni. Clank can sometimes transform into Giant Clank to become a more deadly robot with powerful weaponry. Clank has also wielded the Chronoscepter to manipulate time and access the Netherverse. Clank also at one point starred in a holovision show, Secret Agent Clank.

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